last update 12.IV.2009

Type specimens of mayflies (Ephemeroptera) in the 
Zoological Institute 
of Russian Academy of Sciences 
(in Saint-Petersburg, Russia)

By N.J. Kluge

For details about old type-specimens (original labels, some morphological characters, comments on lectotype designation and synonymy) see:

In 1993-1994 all type specimens of extant mayflies formerly deposited in Moscow State University and Paleontological Institute (Moscow) were transferred to Zoological Institute (S.-Petersburg). Now type collection of the Zoological Institute includes deposited type specimens of extant Ephemeroptera species described by A.D. Hummel (1825), L. Navas (1912), P. Esben-Petersen (1916), O.A. Tshernova and her co-authors (1928-1985), K.A. Brodsky (1930), A.T. Buldovsky (1935), R.S. Kazlauskas (1959-1964), O.Ya. Bajkova (1962-1980), N.D. Sinitshenkova and her co-authors (since 1973 and later), L.A. Kustareva (1976-1984), V.V. Belov (1979, 1981), N.J. Kluge and his co-authors (since 1979 and later), E.A. Novikova and her co-authors (since 1987 and later) and some others. Some type specimens are being temporally held on the premises of the Department of Entomology of St. Petersburg State University.

Type-specimens of mayflies described by T.M. Tiunova and I.M. Levanidova (since 1989 and later) are deposited in the Institute of Biology and Soil Science (Vladivostok). 

All type-specimens of fossil Ephemeroptera described in the USSR and Russia are deposited in the Paleontological Institute (Moscow).

Abbreviations: 
ZIN - Zoological Institute (S-Petersburg) 
L-S-I - imago reared from larva, with larval and subimaginal exuvia; 
L-S - subimago reared from larva, with larval exuvia;
L/S - mature larva with extracted parts of subimago;
- link to the catalogue "Ephemeroptera of the World".

abnormis Tshernova 1949: 139 [Heptagenia] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi 
Subjective synonym (Flowers 1986): lyriformis McDunnough 1924 [Ecdyonurus]

abracadabrus Kluge 1983: 31 [Ecdyonurus] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

acinaciger Kluge 1983: 70 [Baetis] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

albicauda Kluge 1987: 318 [Heptagenia (Heptagenia) sulphurea, subsp.] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi 

albisternum Novikova in Tshernova et al. 1986: 131 [Cloeoptilum] 
Type deposition. ZIN: holotypus (male L-S-I) & paratypi (see Novikova 1987)

alcarrazae Kluge 1992: 18 [Baetis (Caribaetis)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi 

alexandrae Brodsky 1930: 697 [Ameletus] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

alexandri Kluge et Tiunova 1989: 12 [Epeorus (Iron)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi 

altaica Tshernova 1949: 142 [Cinygmula] 
Type deposition. ZIN: syntypi (larvae) 
Subjective synonym (Kluge 1988): cava Ulmer 1927 [Cinygma]

altaicus Kluge 2007: 251 [Ameletus]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

altioculus Kluge 1984: 722 [Choroterpes (Euthraulus)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

amurensis Kluge in Tshernova et al. 1986: 141 [Caenis] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi (see Kluge 1987)

amurensis Navas 1912: 414 [Ephemera] 
Type deposition. ZIN: lectotypus ( subimago, designated by Kluge 1995) & paratypi 
Subjective synonym (Tshernova 1973): orientalis McLachlan 1875 [Ephemera]

anatolii Sinitshenkova 1981: 814 [Epeorus (s.str.)] 
Type deposition. ZIN: holotypus (larva) & paratypus

andrianovae Tshernova et Belov 1982: 283 [Cinygmula] 
Type deposition. ZIN: holotypus ( imago) & paratypi 
Subjective synonym (Belov 1983): malaisei Ulmer 1927 [Cinygmula]

arlecchino Kluge 2007: 254 [Ameletus montanus, subsp.]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

arsenjevi Tshernova 1952: 255 [Heptagenia] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi 
Subjective synonym (Bajkova 1988): flava Rostock 1878 [Heptagenia]

asiatica Tshernova et Bajkova 1960: 413 [Protobehningia] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paratypi

asiatica Sinitshenkova 1973: 13 [Rhithrogena] 
Type deposition. ZIN: holotypus (larva) & paratypi 
Subjective synonym (Kluge 1988): stackelbergi Sinitshenkova 1973 [Rhithrogena]

aspersus Kluge 1980: 569 [Ecdyonurus] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

aurarius Kluge 1983: 29 [Ecdyonurus] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

bacillus Kluge 1983: 72 [Baetis gracilis, subsp.] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

baconaoi Kluge 1994: 276 [Hagenulus (Careospina)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

bajkovae Kluge in Tshernova et al. 1986: 117 [Ecdyonurus] 
Objective synonym: zhiltzovae Bajkova 1975 [Paracinygmula]

balticus Kazlauskas et Sanvaitite 1962: 41 [Baetopus] 
TYPES LOST 
Subjective synonym (Illies 1978): tenellum Albarda 1878 [Centroptilum]

binerve Kluge 1987: 775 [Rhithrogena] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

binotata Sinitshenkova 1982: 65 [Rhithrogena] 
Type deposition. ZIN: holotypus ( imago) & paratypi 
Subjective synonym (Kluge 1995): lepnevae Brodsky 1930 [Rhithrogena]

bipunctata Esben-Petersen 1916: 7 [Heptagenia] 
TYPES LOST 
Subjective synonym (Kluge 1995): flava Rostock 1878 [Heptagenia]

borysthenica Tshernova 1937: 5 [Oligoneuriella] 
Type deposition. ZIN: syntypi (larvae)

brodskyi Kustareva 1976: 66 [Rhithrogena] 
Type deposition. ZIN: holotypus (parts of imago) & paratypus

brodskyi Bajkova 1979: 308 [Siphlonurus] 
Type deposition. ZIN: holotypus (L-S) 
Subjective synonym (Kluge 1985): palaearcticus Tshernova 1930 [Oniscigaster]

caucasica Tshernova 1938: 58 [Cinygma] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi

caucasica Tshernova 1930: 216 [Habroleptoides] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paratypi

cedrensis Sinitshenkova 1977: 44 [Ameletus] 
Type deposition. ZIN: holotypus ( imago) & paratypi

chankae Tshernova 1952: 263 [Siphlonurus] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1985) & paralectotypus

chantauensis Kluge 1981: 1418 [Baetis] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

chimaera Kluge 2008: 398 [Hermanella (Hylister)]
Type deposition. ZIN: holotypus (male L-S-I) & paratypi

citrina Hummel 1825: 71 [Ephemera] (nomen oblitum) 
Type deposition. ZIN: lectotypus ( subimago, designated by Kluge 1995) 
Subjective synonym (Kluge 1995): flava Rostock 1878 [Heptagenia]

confinis Tshernova 1928: 322 [Ecdyonurus] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi 
Subjective synonym (Tshernova 1936): fuscogrisea Retzius 1783 [Ephemera]

corniger Kluge 1991: 16 [Brachycercus] 
Type deposition. ZIN: holotypus (L-S)

cornuta Tshernova 1952: 286 [Ordella] 
Type deposition. ZIN: lectotypus ( larva, designated by Kluge 1995) & paralectotypi

cothurnata Tshernova 1952: 267 [Paraleptophlebia] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paratypi 
Subjective synonym (Bajkova 1976): chocolata Imanishi 1937 [Leptophlebia]

cristatus Bajkova 1976: 586 [Ameletus] 
TYPES LOST

cubensis Malzacher & Naranjo & Gonzalez-Lazo & Kluge 2007 [Caenis]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

cubensis Kluge et Naranjo 1990: 576 [Tricorythodes] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

daedaleus Sinitshenkova 1981: 816 [Epeorus (Belovius)] 
Type deposition. ZIN: holotypus (larva) & paratypi

decolorata Sinitshenkova 1973: 16 [Rhithrogena] 
Type deposition. ZIN: holotypus ( imago) & paratypi

dentata Bajkova 1967: 327 [Ephemerella] 
Type deposition. ZIN: holotypus (larva)

desertus Novikova et Kluge 1987: 14 [Baetis (Labiobaetis)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

diversicolor Tshernova 1952: 266 [Baetis] 
TYPES LOST

dracon Kluge 1983: 27 [Ecdyonurus] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

eatoni Brodsky 1930: 33 [Ametropus]
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypus
Subjective synonym: fragilis Albarda 1878 [Ametropus]

ermolenkoi Tshernova 1981: 327 [Epeorus (Belovius)] 
Type deposition. ZIN: holotypus ( imago) & paratypi

eugeniae Kluge 1983: 1116 [Rhithrogena] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

eugenii Sinitshenkova et Varychanova 1989: 576 [Ameletus] 
Type deposition. ZIN: holotypus ( imago) & paratypi 
Subjective synonym (Kluge 1995): inopinatus Eaton 1887 [Ameletus]

europaeus Kluge 1991: 19 [Brachycercus] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

evanescens Kluge 1994: 276 [Hagenulus (Careospina)] 
Type deposition. ZIN: holotypus ( imago) & paratypi

excisa Sinitshenkova 1979: 7 [Rhithrogena] 
Type deposition. ZIN: holotypus ( imago) & paratypi

feles Kluge 1980: 562 [Baetis] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

femorata Tshernova 1972: 611 [Asiatella] 
Type deposition. ZIN: holotypus (larva)

fenestratum Kazlauskas 1963: 588 [Pseudocloeon] 
Type deposition. ZIN: holotypus (parts of larva)

frater Tshernova 1938: 58 [Ecdyonurus] 
Type deposition. ZIN: holotypus ( imago) 
Subjective synonym (Thomas & Dia 1982): znojkoi Tshernova 1938 [Ecdyonurus]

frolenkoi Sinitshenkova 1981: 815 [Epeorus (Belovius)] 
Type deposition. ZIN: holotypus (larva) & paratypi

fuscus Sinitshenkova 1976: 859 [Iron] 
Type deposition. ZIN: holotypus (larva) & paratypi

gnom Kluge 1983: 74 [Pseudocloeon] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

gornostajevi Tshernova 1981: 327 [Epeorus (Belovius)] 
Type deposition. ZIN: holotypus ( imago) & paratypi

gracilis Tshernova 1952: 257 [Ephemerella] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

grallator Kluge et Naranjo 1990: 575 [Tricorythodes] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

grandifolia Tshernova 1952: 256 [Cinygmula] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

grandis Kluge 1994: 265 [Hagenulus (Turquinophlebia)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

grisea Navas 1912: 416 [Andromina] 
Type deposition. ZIN: syntypi ( subimagines)

grunini Sinitshenkova 1982: 59 [Iron] 
Type deposition. ZIN: holotypus ( imago) & paratypi 
Subjective synonym (Kluge & Tiunova 1989): aesculus Imanishi 1934 [Epeorus]

gultsha Kustareva 1984: 16 [Iron] 
TYPE LOST

guranica Belov 1981: 601 [Heptagenia] 
Type deposition. ZIN: holotypus ( imago) & paratypi

gurvitshi Kustareva 1984: 20 [Iron] 
TYPES LOST

heptapotamicus Brodsky 1930: 693 [Baetis] 
Type deposition. ZIN: holotypus ( imago)

heterophillum Kluge et Novikova 1992: 74 [Cloeon (Procloeon)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

hissari Kluge 1985: 1586 [Caenis] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

hissaricus Novikova 1991: 793 [Baetis ursinus, subsp.] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

hyalinus Esben-Petersen 1916: 6 [Ecdyonurus] nom. praeocc. 
Type deposition. ZIN: holotypus ( imago) 
Objective synonym: peterseni Lestage 1930 [Ecdyonurus] 
Subjective synonym (Flowers 1986): lyriformis McDunnough 1924 [Ecdyonurus]

imanica Bajkova 1972: 207 [Rhithrogena] 
Type deposition. ZIN: holotypus ( imago) 
Subjective synonyms (Kluge 1980, Flowers 1986): werestschagini Tshernova 1952 [Heptagenia]; 
simplicioides McDunnough 1924 [Heptagenia]

immanis Kluge 1985: 18 [Siphlonurus] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

inaccessibile Kluge et Naranjo 1994: 778 [Euthyplocia] 
Type deposition. ZIN: holotypus ( imago) & paratypi

inexpectatum Tshernova 1928: 320 [Pseudocloeon] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi 

inferior Kluge 1991: 133 [Cloeodes] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

intermedia Brodsky 1930: 708 [Iron montanus, var.] 
Type deposition. ZIN: syntypi (imagines)

inversus Kluge 1980: 569 [Ecdyonurus] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

irina Tshernova et Belov 1982: 286 [Cinygmula] 
Type deposition. ZIN: paratypus ( imago); holotypus lost

issyksuvensis Brodsky 1930: 688 [Baetis] 
Type deposition. ZIN: syntypi

itayana Kluge 2007: 130 [Oligineuria]
Type deposition. ZIN: holotypus (L-S/I) & paratypi

karasuensis Kustareva 1976: 58 [Drunella] 
Type deposition. ZIN: holotypus (larva) 
Subjective synonym (Kluge 1995): ignita Poda 1761 [Ephemera]

kazlauskasi Kluge 1983: 78 [Centroptilum] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

kirgisicus Kustareva 1984: 14 [Iron] 
TYPES LOST

kogistani Novikova et Kluge 1994: 637 [Baetis (Nigrobaetis)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

kopetdagi Kluge 1985: 1586 [Caenis] 
Type deposition. ZIN: holotypus ( L-S-I) & paratypi

kozhovi Bajkova 1967: 327 [Ephemerella] 
Type deposition. ZIN: holotypus ( imago) & paratypi

kurenzovi Bajkova 1965: 301 [Cinygmula] 
Type deposition. ZIN: holotypus ( imago) & paratypi

labiatus Sinitshenkova 1981: 75 [Ameletus] 
Type deposition. ZIN: holotypus (larva) & paratypi 

laciniosa Sinitshenkova 1979: 816 [Rhithrogena] 
Type deposition. ZIN: holotypus (larva) & paratypi

latericius Sinitshenkova 1982: 53 [Iron] 
Type deposition. ZIN: holotypus ( imago) & paratypi 
Subjective synonym (Kluge & Tiunova 1989): maculatus Tshernova 1949 [Iron]

latifrons Tshernova et Belov 1982: 287 [Cinygmula] 
Type deposition. ZIN: paratypi; holotypus lost
Subjective synonym (Kluge 1995): hirasana Imanishi 1935 [Cinygma]

latipes Tshernova 1952: 273 [Ephemerella] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi 
Subjective synonym (Tiunova & Belov 1984): cryptomeria Imanishi 1937 [Ephemerella]

lenoki Tshernova 1952: 275 [Ephemerella] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

lepnevae Tshernova 1949: 152 [Ephemerella] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

lepnevae Brodsky 1930: 39 [Rhithrogena] 
Type deposition. ZIN: syntypi ( imagines)

levanidovae Tshernova 1952: 274 [Ephemerella] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

levanidovae Sinitshenkova 1982: 60 [Iron] 
Type deposition. ZIN: holotypus ( imago) & paratypi
Subjective synonym (Kluge & Tiunova 1989): aesculus Imanishi 1934 [Epeorus]

levanidovi Tshernova et Belov 1982: 289 [Cinygmula] 
Type deposition. ZIN: holotypus ( imago)

levis Navas 1912: 415 [Epeorus] 
Type deposition. ZIN: holotypus ( imago)

lilliputian Kluge 1991: 134 [Paracloeodes] 
Type deposition. ZIN: holotypus (L-S-I) & paratypus 

lithuanica Kazlauskas 1959: 168 [Ephemerella] 
TYPES LOST 
Subjective synonym (Illies 1978): karelica Tiensuu 1935 [Eurylophella]

longifolius Kluge 1992: 44 [Baetis (Fallceon)] 
Type deposition. ZIN: holotypus (male L-S-I) & paratypi

longilobata Tshernova 1928: 113 [Leptophlebia (Paraleptophlebia)] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paratypi

longipes Tshernova 1952: 274 [Ephemerella] 
Type deposition. ZIN: holotypus (larva)
Subjective synonym (Tshernova et al. 1986): lepnevae Tshernova 1949 [Ephemerella]

longulus Sinitshenkova 1981: 77 [Ameletus] 
Type deposition. ZIN: holotypus (larva) & paratypi

lunata Tshernova 1952: 269 [Paraleptophlebia] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi 
Subjective synonym (Kluge 1995): strandii Eaton 1901 [Leptophlebia]

lutea Kluge 1987: 311 [Heptagenia (Heptagenia)] nom. praeocc. 
Type deposition. ZIN: holotypus (L-S-I) & paratypi
Subjective synonym (Sartori 1992): samochai Demoulin 1973 [Sigmoneuria]

macronyx Kluge in Tshernova et al. 1986: 141 [Caenis] 
Type deposition. ZIN: holotypus (mature larva) & paratypi

macronyx Kluge et Novikova 1992: 76 [Cloeon (Procloeon)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

macronyx Kluge 1996 [Metreplecton]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

maculata Tshernova1952: 287 [Ordella] 
Type deposition. ZIN: syntypi (larvae)

maculatus Tshernova 1949: 148 [Iron] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge & Tiunova 1989) & paralectotypi

magnus Tshernova 1952: 284 [Brachycercus] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1991) & paralectotypi
Subjective synonym (Kluge 1991): harrisella Curtis 1834 [Brachycercus]

maritimus Kluge 1983: 76 [Centroptilum] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

markevitshi Belov 1979: 577 [Uracanthella] 
Type deposition. ZIN: holotypus (larva) & paratypi
Subjective synonym: lenoki Tshernova 1952 [Ephemerella]

micracantha Kluge 1989: 4 [Heptagenia (Dacnogenia) coerulans, subsp.] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

micus Bajkova 1976: 583 [Ameletus] 
Type deposition. ZIN: holotypus (larva)

miliaria Tshernova 1952: 286 [Ordella] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

minima Sinitshenkova 1973: 13 [Rhithrogena] 
Type deposition. ZIN: holotypus (larva) & paratypi

minuta Brodsky 1930: 709 [Iron montanus, var.] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi

minutus Tshernova 1952: 285 [Brachycercus] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1991) & paralectotypi

modesta Brodsky 1930: 31 [Ephemera] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paratypi
Subjective synonym (Tshernova 1973): orientalis McLachlan 1875 [Ephemera]

mongolica Bajkova et Varychanova 1978: 14 [Heptagenia] 
TYPES LOST
Subjective synonym (Braasch 1983): joernensis Bengtsson 1909 [Ecdyonurus]

mongolicus Tshernova 1952: 266 [Baetis] 
TYPES LOST

montana Brodsky 1930: 704 [Iron montanus, var.] 
Type deposition. ZIN: syntypi

montanus Kluge et Naranjo 1990: 574 [Tricorythodes] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

mycetopis Brodsky 1930: 691 [Baetis] 
Type deposition. ZIN: holotypus ( imago)

naranjoi Kluge 1992: 19 [Baetis (Americabaetis)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

natans Buldovsky 1935: 153 [Chankagenesia] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paratypi

nigridorsum Tshernova 1934: 240 [Eopolymitarcys] 
Type deposition. ZIN: syntypi (imagos)

nigripilosus Sinitshenkova 1976: 860 [Iron] 
Type deposition. ZIN: holotypus (larva) & paratypi

nigromaculata Brodsky 1930: 708 [Iron montanus, var.] 
Type deposition. ZIN: syntypi

nikolskyi Tshernova 1948: 1453 [Acanthametropus] 
Type deposition. ZIN: holotypus (larva)

ninae Kluge 1995 [Epeorus (Belovius)] 
Type deposition. ZIN: holotypus ( imago) & paratypi 
Objective synonym: sinitshenkovae Tshernova 1981 [Epeorus (Belovius)]

nuda Tshernova 1949: 53 [Ephemerella] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

obscuriventris Tshernova 1952: 266 [Baetis] 
TYPES LOST

obtusiceps Tshernova 1952: 265 [Baetis] 
TYPES LOST

orbiticola Kluge 1987: 305 [Heptagenia (Kageronia)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

oreophila Kustareva 1978: 92 [Cinygmula] 
Type deposition. ZIN: holotypus ( imago) & paratypi

oreophilus Kluge 1982: 15 [Baetis] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

oriens Bae et McCafferty 1991: 54 [Potamanthus luteus, subsp.] 
Type deposition. ZIN: holotypus (L-S-I)

orientalis Kluge 1983: 74 [Baetis atrebatinus, subsp.] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi
Subjective synonym (Kluge 1995): atrebatinus Eaton 1870 [Baetis]

orientalis Tshernova 1952: 279 [Ephemerella] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi
Subjective synonym (Tshernova et al. 1986): levanidovae Tshernova 1952 [Ephemerella]

oriente Kluge 1994: 282 [Hagenulus (Traverina)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

ornata Tshernova 1972: 613 [Ephemerellina] 
Type deposition. ZIN: holotypus ( subimago) & paratypus

ornatipennis Tshernova 1938: 55 [Ecdyonurus] 
Type deposition. ZIN: holotypus ( imago)

ornatum Tshernova 1928: 319 [Procloeon] 
Type depositions. ZIN: paralectotypi; 
Zool. Mus. Hamburg Univ.: lectotypus ( imago), designated by Sowa 1975: 108)
Subjective synonym (Jacob 1974): bifidum Bengtsson 1912 [Cloeon]

pacoi Kluge 1994: 269 [Hagenulus (Poecilophlebia)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

padunica Kazlauskas 1963: 584 [Ephemerella (Tortleya)] 
TYPES LOST

palaearcticus Tshernova 1930: 217 [Oniscigaster] 
Type deposition. ZIN: holotypus ( imago)

pallidus Tshernova 1928: 114 [Brachycercus] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1991) & paralectotypi
Subjective synonym (Kluge 1991): harrisella Curtis 1834 [Brachycercus]

pamirensis Kustareva 1984: 20 [Iron] 
TYPE LOST

paradoxa Buldovsky 1935: 157 [Anagenesia] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paratypi

parvus Kluge 1979: 807 [Ameletus] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

pellucida Brodsky 1930: 35 [Cinygma] 
Type deposition. ZIN: holotypus ( imago)

perflava Brodsky 1930: 701 [Heptagenia] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1987) & paralectotypus

peterseni Lestage 1930: 199 [Ecdyonurus] 
Type deposition. ZIN: holotypus ( imago)
Invalid objective synonym: hyalinus Esben-Petersen 1916 [Ecdyonurus] nom. praeocc.
Subjective synonym (Flowers 1986): lyriformis McDunnough 1924 [Ecdyonurus]

petropolitanum Kluge et Novikova 1992: 68 [Cloeon (Intercloeon)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

petrovi Tshernova 1938: 55 [Baetis] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypus

pilatus Sinitshenkova 1981: 73 [Ameletus] 
Type deposition. ZIN: holotypus (larva) & paratypi
Subjective synonym (Kluge 1995): camtshaticus Ulmer 1923 [Ameletus]

planifrons Kluge 1992: 15 [Baetis (Caribaetis)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

platysma Tshernova 1941: 239 [Heptagenia] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi
Subjective synonym (Kluge 1987): perflava Brodsky 1930 [Heptagenia]

plesius Kluge 2008: 394 [Simothraulopsis (Maculognathus)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

polita Bajkova 1970: 146 [Isonychia] 
Type deposition. ZIN: holotypus ( imago)
Subjective synonym (Tshernova et al. 1986): nikolskyi Tshernova 1948 [Acanthametropus]

pontica Kluge 1984: 40 [Habroleptoides] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

procerus Bajkova 1976: 584 [Ameletus] 
Type deposition. ZIN: holotypus (larval exuvia) & paratypi (see Kluge 1995)
Subjective synonym (Kluge 1995): montanus Imanishi 1930 [Ameletus]

pseudothermicus Kluge 1983: 65 [Baetis] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

putoranica Kluge 1980: 576 [Cinygmula] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

quadrata Kluge & Zhou (in Kluge & Zhou & Jacobus & McCafferty) 2004 [Ephemerella]
Type deposition. ZIN: holotypus (larva) & paratypi

quadrinotata Sinitshenkova 1982: 63 [Rhithrogena] 
Type deposition. ZIN: holotypus ( imago) & paratypi
Subjective synonym (Kluge 1995): bajkovae Sowa 1973 [Rhithrogena]

quadripunctata Kluge 1989: 4 [Heptagenia (Heptagenia)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

rautiani Sinitshenkova 1982: 53 [Epeorus (Epeorus)] 
Type deposition. ZIN: paratypi (larvae, subimageos, imageos); holotypus lost 

rheophilus Brodsky 1930: 712 [Iron] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

rohdendorfi Tshernova 1985: 15 [Rhoenanthus] 
Type deposition. ZIN: holotypus ( imago)
Subjective synonym (Bae & McCafferty 1991): coreanus Yoon et Bae 1985 [Potamanthus]

romantzovi Kluge 1988: 76 [Ephemera] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

rossicus Kluge 2007: 253 [Ameletus montanus]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

rossicus Tshernova 1928: 321 [Ecdyonurus] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi
Subjective synonym (Kluge 1987): fuscogrisea Retzius 1783 [Ephemera]

rubrofasciatus Brodsky 1930: 716 [Ecdyonurus] 
Type deposition. ZIN: holotypus ( imago)

sabalo Kluge 2008: 389 [Simothraulopsis (Maculognathus)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

sacculobranchis Kluge et Naranjo 1990: 570 [Tricorythodes] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

scalaris Kluge 1983: 34 [Ecdyonurus] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

setigera Bajkova 1965: 327 [Ephemerella] 
Type deposition. ZIN: holotypus (larva)

sextus Kluge 1992: 45 [Baetis (Fallceon)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

sextus Kluge 1994: 261 [Hagenulus (Borinquena)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

shadini Kazlauskas 1964: 170 [Pseudocentroptilum] 
Type deposition. ZIN: lectotypus (parts of larva, designated by Kluge 1995) & paralectotypus
Subjective synonym (Kluge & Novikova 1992): nana Bogoescu 1951 [Centroptilum]

sibirica Tshernova 1952: 278 [Ephemerella] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi
Subjective synonym (Bajkova 1972): ignita Poda 1761 [Ephemera]

sibirica Tiunova & Kluge & Ishiwata 2004 [Isonychia ussurica] 
Type deposition. ZIN: holotypus (L-S-I) and paratypi

sibirica Brodsky 1930: 37 [Rhithrogena] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi

sibiricum Kazlauskas 1963: 589 [Pseudocloeon] 
Type deposition. ZIN: holotypus (parts of larva) & paratypi

sierramaestrae Kluge 1994: 274 [Hagenulus (Careospina) hespera, subsp.] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

sierramaestrae Kluge et Naranjo 1990: 570 [Tricorythodes] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

silvaticus Kluge 1983: 67 [Baetis] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

sinitshenkovae Tshernova 1981: 332 [Epeorus (Belovius)] 
Type deposition. ZIN: holotypus ( imago) & paratypi
Objective synonym: ninae Kluge 1995 [Epeorus (Belovius)]

sinitshenkovae Tiunova & Kluge & Ishiwata 2004: 28 [Isonychia vshivkovae] 
Type deposition. ZIN: holotypus (L-S-I) and paratypi

smirnovi Tshernova 1978: 74 [Cinygmula] 
Type deposition. ZIN: holotypus ( imago) & paratypi
Subjective synonym (Kluge 1995): pellucida Brodsky 1930: 35 [Cinygma]

soldatovi Tshernova 1952: 253 [Heptagenia] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1987) & paralectotypi 
Subjective synonym (Bajkova 1972, Kluge 1987): sulphurea Muller 1776 [Ephemera]

solida Bajkova 1980: 796 [Ephemerella] 
TYPES LOST

spiniventre Kluge et Novikova 1992: 65 [Cloeon (Intercloeon)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

spinosa Tshernova et Belov 1982: 291 [Cinygmula] 
Type deposition. ZIN: holotypus ( imago)
Subjective synonym (Tshernova et al. 1986): levanidovi Tshernova et Belov 1982 [Cinygmula]

stackelbergi Sinitshenkova 1973: 18 [Rhithrogena] 
Type deposition. ZIN: holotypus ( imago) & paratypus

stipposus Kluge 1982: 18 [Baetis] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi
Subjective synonym (Godunko & al. 2004): braaschi Zimmermann 1980 [Baetis]

sublongicauda Tshernova 1949: 306 [Palingenia] 
Type deposition. ZIN: syntypi

submontana Brodsky 1930: 683 [Ephemerella] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

sumbarensis Kluge 1984: 722 [Choroterpes (Euthraulus)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

superior Kluge 1991: 130 [Cloeodes] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

tadjikistanicum Brodsky 1930: 696 [Cloeon] 
Type deposition. ZIN: syntypi ( imagines)

tadzhikorum Tshernova 1930: 218 [Heptagenia] 
Type deposition. ZIN: holotypus ( imago)
Subjective synonym (Kluge 1987): perflava Brodsky 1930 [Heptagenia]

taeniata Tshernova 1952: 279 [Ephemerella] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi
Subjective synonym (Bajkova 1972): aurivillii Bengtsson 1908 [Chitonophora]

talasi Novikova et Kluge 1994: 635 [Baetis (Nigrobaetis)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

tenax Tshernova 1952: 273 [Ephemerella] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi
Subjective synonym (Kluge 1995): triacantha Tshernova 1949 [Ephemerella]

testudineus Kluge 1992: 45 [Baetis (Fallceon)] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

thani Tshernova 1972: 609 [Vietnamella] 
Type deposition. ZIN: holotypus (larva) & paratypus

thymalli Tshernova 1952: 277 [Ephemerella] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi
Subjective synonym (Kluge 1995): nuda Tshernova 1949 [Ephemerella]

tianshanica Kustareva 1984: 18 [Iron] 
TYPES LOST

tianshanica Brodsky 1930: 37 [Rhithrogena] 
Type deposition. ZIN: lectotypus (L/S, designated by Kluge 1995) & paralectotypi

tianshanica Kustareva 1984: 26 [Stenonema] 
Type deposition. ZIN: holotypus ( imago)
Subjective synonym (Kluge 1995): rubrofasciatus Brodsky 1930 [Ecdyonurus]

transbajkalica Tshernova 1973: 333 [Ephemera] 
Type deposition. ZIN: holotypus ( imago) & paratypi

transiliensis Brodsky 1930: 689 [Baetis] 
Type deposition. ZIN: holotypus ( imago)

triacantha Tshernova 1949: 41 [Ephemerella] 
TYPES LOST

tricolor Tshernova 1928: 320 [Baetis] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi

tshernovae Edmunds et Traver 1959 [Behningia] 
Type deposition. ZIN: holotypus ( imago) & paratypi

tshernovae Bajkova 1962: 202 [Ephemerella] 
Type deposition. ZIN: holotypus (larva)

tshernovae Sinitshenkova 1982: 56 [Iron] 
Type deposition. ZIN: holotypus ( imago) & paratypi 
Subjective synonym (Kluge & Tiunova 1989): maculatus Tshernova 1949 [Iron]

tshernovae Kazlauskas 1963: 582 [Neoephemera] 
Type deposition. ZIN: holotypus (larva) & paratypi

tuberculatum Kazlauskas 1963: 586 [Pseudocloeon (Baetiella)] 
Type deposition. ZIN: holotypus (parts of larva)

tubulatus Tshernova 1952: 285 [Brachycercus] 
Type deposition. ZIN: holotypus (larva)

ulmeri Brodsky 1930: 686 [Caenis] 
Type deposition. ZIN: syntypi ( imagoes) 
Subjective synonym (Malzacher 1993): robusta Eaton 1884 [Caenis]

unguiculatum Tshernova 1941: 242 [Cloeon] 
Type deposition. ZIN: lectotypus (parts of larva, designated by Kluge 1996)

unicolor Tshernova 1952: 250 [Rhithrogena] 
Type deposition. ZIN: lectotypus (L/S, designated by Kluge 1995) & paralectotypi
Subjective synonym (Kluge 1995): lepnevae Brodsky 1930 [Rhithrogena]

unicolorata Tshernova 1979: 9 [Cinygmula] 
Type deposition. ZIN: holotypus ( imago) & paratypi

ursinus Kazlauskas 1963: 585 [Baetis] 
Type deposition. ZIN: holotypus (parts of larva)

ussurica Bajkova 1970: 146 [Isonychia] 
Type deposition. ZIN: holotypus ( imago) & paratypus

ussuricus Kluge 1983a: 68 [Baetis] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

vernalis Tshernova 1928: 115 [Siphlonurus] 
Type deposition. ZIN: holotypus (genitals of imago)
Subjective synonym (Puthz 1977): aestivalis Eaton 1903 [Siphlurus]

verrucosa Kluge 1980: 566 [Ephemerella] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi
Subjective synonym (Kluge 1995): nuda Tshernova 1949 [Ephemerella]

vladivostokica Kluge 1982: 112 [Paraleptophlebia] 
Type deposition. ZIN: holotypus (L-S-I) & paratypi

vshivkovae Tiunova & Kluge & Ishiwata 2004: 24 [Isonychia] 
Type deposition. ZIN: holotypus (L-S-I) and paratypi

werestschagini Tshernova 1952: 253 [Heptagenia] 
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi
Subjective synonym (Flowers 1986): simplicioides McDunnough 1924 [Heptagenia]

zachvatkini Tshernova 1952: 257 [Cinygmula] 
Type deposition. ZIN: holotypus ( imago) & paratypi
Subjective synonym (Tshernova et al. 1986): levis Navas 1912 [Epeorus]

zaitsevi Tshernova 1981: 326 [Epeorus (Epeorus)] 
Type deposition. ZIN: holotypus ( imago)

zapekinae Bajkova 1967: 327 [Ephemerella] 
Type deposition. ZIN: holotypus ( imago) & paratypi

zhelochovtsevi Tshernova 1952: 263 [Siphlonurus] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1985a) & paralectotypus

zhiltzovae Tshernova 1972: 605 [Ecdyonurus] 
Type deposition. ZIN: holotypus ( imago) & paratypi

zhiltzovae Bajkova 1975: 54 [Paracinygmula] 
Type deposition. ZIN: holotypus (larva) & paratypi
Objective synonym: bajkovae Kluge 1986 [Ecdyonurus]

zhiltzovae Sinitshenkova 1979: 811 [Rhithrogena] 
Type deposition. ZIN: holotypus ( imago) & paratypi

zimini Tshernova 1930: 214 [Cloeon] 
Type deposition. ZIN: holotypus ( imago) & paratypi
Subjective synonym (Kluge 1995): diptera Linnaeus 1761 [Ephemera]

znojkoi Tshernova 1938: 56 [Ecdyonurus] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi

znojkoi Tshernova 1938: 59 [Iron] 
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi